characters of a species—and so few even among naturalists have ever compared, accurately, considerable numbers of individuals, that the conception of variability as a general characteristic of all dominant and widespread species, large in its amount and affecting, not a few, but considerable masses of the individuals which make up the species, will be to many entirely new. Equally important is the fact that the variability extends to every organ and every structure, external and internal; while perhaps most important of all is the independent variability of these several parts, each one varying without any constant or even usual dependence on, or correlation with, other parts. No doubt there is some such correlation in the differences that exist between species and species—more developed wings usually accompanying smaller feet and vice versâ—but this is, generally, a useful adaptation which has been brought about by natural selection, and does not apply to the individual variability which occurs within the species.
It is because these facts of variation are so important and so little understood, that they have been discussed in what will seem to some readers wearisome and unnecessary detail. Many naturalists, however, will hold that even more evidence is required; and more, to almost any amount, could easily have been given. The character and variety of that already adduced will, however, I trust, convince most readers that the facts are as stated; while they have been drawn from a sufficiently wide area to indicate a general principle throughout nature.
If, now, we fully realise these facts of variation, along with those of rapid multiplication and the struggle for existence, most of the difficulties in the way of comprehending how species have originated through natural selection will disappear. For whenever, through changes of climate, or of altitude, or of the nature of the soil, or of the area of the country, any species are exposed to new dangers, and have to maintain themselves and provide for the safety of their offspring under new and more arduous conditions, then, in the variability of all parts, organs, and structures, no less than of habits and intelligence, we have the means of producing modifications which will certainly bring the species into harmony with its
