Page:Darwinism by Alfred Wallace 1889.djvu/378

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354
DARWINISM
CHAP.

for being carried by mammalia or birds, and for floating in the water, or through the air, while many are so small and so light that there is practically no limit to the distances they may be carried by gales and hurricanes.

We may, therefore, feel quite certain that the means of distribution that have enabled the larger mammalia to reach the most remote regions from a common starting-point, will be at least as efficacious, and usually far more efficacious, with all other land animals and plants; and if in every case the existing distribution of this class can be explained on the theory of oceanic and continental permanence, with the limited changes of sea and land already referred to, no valid objections can be taken against this theory founded on anomalies of distribution in other orders. Yet nothing is more common than for students of this or that group to assort that the theory of oceanic permanence is quite inconsistent with the distribution of its various species and genera. Because a few Indian genera and closely allied species of birds are found in Madagascar, a land termed "Lemuria" has been supposed to have united the two countries during a comparatively recent geological epoch; while the similarity of fossil plants and reptiles, from the Permian and Miocene formations of India and South Africa, has been adduced as further evidence of this connection. But there are also genera of snakes, of insects, and of plants, common to Madagascar and South America only, which have been held to necessitate a direct land connection between these countries. These views evidently refute themselves, because any such land connections must have led to a far greater similarity in the productions of the several countries than actually exists, and would besides render altogether inexplicable the absence of all the chief types of African and Indian mammalia from Madagascar, and its marvellous individuality in every department of the organic world.[1]

Powers of Dispersal as illustrated by Insular Organisms.

Having arrived at the conclusion that our existing oceans have remained practically unaltered throughout the Tertiary and Secondary periods of geology, and that the distribution of the

  1. For a full discussion of this question, see Island Life, pp. 390-420.