leeward side of an atoll, there may be one or more navigable
passages leading from the lagoon to the open sea.
Though corals flourish everywhere under suitable conditions in tropical seas, coral reefs and atolls are by no means universal in the torrid zone. The Atlantic Ocean is remarkably free from coral formations, though there are numerous reefs in the West Indian islands, off the south coast of Florida, and on the coast of Brazil. The Bermudas also are coral formations, their high land being formed by sand accumulated by the wind and cemented into rock, and are remarkable for being the farthest removed from the equator of any recent reefs, being situated in 32° N. lat. In the Pacific Ocean there is a vast area thickly dotted with coral formations, extending from 5° N. lat. to 25° S. lat., and from 130° E. long, to 145° W. long. There are also extensive reefs in the westernmost islands of the Hawaiian group in about 25° N. lat. In the Indian Ocean, the Laccadive and Maldive islands are large groups of atolls off the west and south-west of India. Still farther south is the Chagos group of atolls, and there are numerous reefs off the north coast of Madagascar, at Mauritius, Bourbon and the Seychelles. The Cocos-Keeling Islands, in 12° S. lat. and 96° E. long., are typical atolls in the eastern part of the Indian Ocean.
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Diagram showing the formation of an atoll during subsidence. |
The remarkable characters of barrier reefs and atolls, their isolated position in the midst of the great oceans the seemingly unfathomable depths from which they rise their peaceful and shallow lagoons and inner channels, their narrow strips of land covered with coco-nut palms and other vegetation, and rising but a few feet above the level of the ocean, naturally attracted the attention of the earlier navigators, who formed sundry speculations as to their origin. The poet-naturalist, A. von Chamisso, was the first to propound a definite theory of the origin of atolls and encircling reefs, attributing their peculiar features to the natural growth of corals and the action of the waves. He pointed out that the larger and more massive species of corals flourish best on the outer sides of a reef, whilst the more interior corals are killed or stunted in growth by the accumulation of coral and other debris. Thus the outer edge of a submerged reef is the first to reach the surface, and a ring of land being formed by materials piled up by the waves, an atoll with a central lagoon is produced. Chamisso’s theory necessarily assumed the existence of a great number of submerged banks reaching nearly, but not quite, to the surface of the sea in the Pacific and Indian oceans, and the difficulty of accounting for the existence of so many of these led C. Darwin to reject his views and bring forward an explanation which may be called the theory of subsidence. Starting from the well-known premise that reef-building species of corals do not flourish in a greater depth of water than 20 fathoms, Darwin argued that all coral islands must have a rocky base, and that it was inconceivable that, in such large tracts of sea as occur in the Pacific and Indian oceans, there should be a vast number of submarine peaks or banks all rising to within 20 or 30 fathoms of the surface and none emerging above it. But on the supposition that the atolls and encircling reefs were formed round land which was undergoing a slow movement of subsidence, their structure could easily be explained. Take the case of an island consisting of a single high peak. At first the coral growth would form a fringing reef clinging to its shores. As the island slowly subsided into the ocean the upward growth of coral would keep the outer rim of the reef level with or within a few fathoms of the surface, so that, as subsidence proceeded, the distance between the outer rim of the reef and the sinking land would continually increase, with the result that a barrier-reef would be formed separated by a wide channel from the central peak. As corals and other organisms with calcareous skeletons live in the channel, their remains, as well as the accumulation of coral and other debris thrown over the outer edge of the reef, would maintain the channel at a shallower depth than that of the ocean outside. Finally, if the subsidence continued, the central peak would disappear beneath the surface, and an atoll would be left consisting of a raised margin of reef surrounding a central lagoon, and any pause during the movement of subsidence would result in the formation of raised islets or a strip of dry land along the margin of the reef. Darwin’s theory was published in 1842, and found almost universal acceptance, both because of its simplicity and its applicability to every known type of coral-reef formation, including such difficult cases as the Great Chagos Bank, a huge submerged atoll in the Indian Ocean.
Darwin’s theory was adopted and strengthened by J. D. Dana, who had made extensive observations among the Pacific coral reefs between 1838 and 1842, but it was not long before it was attacked by other observers. In 1851 Louis Agassiz produced evidence to show that the reefs off the south coast of Florida were not formed during subsidence, and in 1863 Karl Semper showed that in the Pelew islands there is abundant evidence of recent upheaval in a region where both atolls and barrier-reefs exist. Latterly, many instances of recently upraised coral formations have been described by H. B. Guppy, J. S. Gardiner and others, and Alexander Agassiz and Sir J. Murray have brought forward a mass of evidence tending to shake the subsidence theory to its foundations. Murray has pointed out that the deep-sea soundings of the “Tuscarora” and “Challenger” have proved the existence of a large number of submarine elevations rising out of a depth of 2000 fathoms or more to within a few hundred fathoms of the surface. The existence of such banks was unknown to Darwin, and removes his objections to Chamisso’s theory. For although they may at first be too far below the surface for reef-building corals, they afford a habitat for numerous echinoderms, molluscs, crustacea and deep-sea corals, whose skeletons accumulate on their summits, and they further receive a constant rain of the calcareous and silicious skeletons of minute organisms which teem in the waters above. By these agencies the banks are gradually raised to the lowest depth at which reef-building corals can flourish, and once these establish themselves they will grow more rapidly on the periphery of the bank, because they are more favourably situated as regards food-supply. Thus the reef will rise to the surface as an atoll, and the nearer it approaches the surface the more will the corals on the exterior faces be favoured, and the more will those in the centre of the reef decrease, for experiment has shown that the minute pelagic organisms on which corals feed are far less abundant in a lagoon than in the sea outside. Eventually, as the margin of the reef rises to the surface and material is accumulated upon it to form islets or continuous land, the coral growth in the lagoon will be feeble, and the solvent action of sea-water and the scour of the tide will tend to deepen the lagoon. Thus the considerable depth of some lagoons, amounting to 40 or 50 fathoms, may be accounted for. The observations of Guppy in the Solomon islands have gone far to confirm Murray’s conclusions, since he found in the islands of Ugi, Santa Anna and Treasury and Stirling islands unmistakable evidences of a nucleus of volcanic rock, covered with soft earthy bedded deposits several hundred feet thick. These deposits are highly fossiliferous in parts, and contain the remains of pteropods, lamellibranchs and echinoderms, embedded in a foraminiferous deposit mixed with volcanic debris, like the deep-sea muds brought up by
