distinguished by features of structure or habit. Certain extremely aberrant Diptera, which, in consequence of the adoption of a parasitic mode of life, have undergone great structural modification, are further remarkable for their peculiar mode of reproduction, on account of which the families composing the group are often termed Pupipara. In these forms the pregnant female, instead of laying eggs, as Diptera usually do, or even producing a number of minute living larvae, gives birth at one time but to a single larva, which is retained within the oviduct of the mother until adult, and assumes the pupal state immediately on extrusion. The Pupipara are also termed Eproboscidea (although they actually possess a well-developed and functional proboscis), and by some dipterists the Eproboscidea are regarded as a suborder and contrasted as such with the rest of the Diptera, which are styled the suborder Proboscidea. By other writers Proboscidea and Eproboscidea are treated as primary divisions of the Cyclorrhapha. In reality, however, the families designated Eproboscidea (Hippoboscidae, Braulidae, Nycteribiidae and Streblidae), are not entitled to be considered as constituting either a suborder, or even a main division of the Cyclorrhapha; they are simply Cyclorrhapha much modified owing to parasitism, and in view of the closely similar mode of reproduction in the tsetse-flies the special designation Pupipara should be abandoned. Before leaving the subject of classification it may be noted in passing that in 1906 Professor Lameere, of Brussels, proposed a scheme for the classification of Diptera which as regards both the limits of the families and their grouping into higher categories differs considerably from that in current use.
Little light on the relationship and evolution of the various families of Diptera is afforded by fossil forms, since as a rule the latter are readily referable to existing families. With the exception of a few species from the Solenhofen lithographic Oolite, fossil Diptera belong to the Tertiary Period, during which the members of this order attained a high degree of development. In amber, as proved by the deposits on the shores of the Baltic, the proverbial “fly” is more numerous than any other creatures, and with very few exceptions representatives of all the existing families have been found. The famous Tertiary beds at Florissant, Colorado, have yielded a considerable number of remarkably well-preserved Tipulidae (in which family are included the most primitive of existing Diptera), as also species belonging to other families, such as Mycetophilidae and even Oestridae.
Diptera as an order are probably more widely distributed over the earth’s surface than are the representatives of any similar division of the animal kingdom. Flies seem capable of adapting themselves to extremes of cold equally as well as to those of heat, and species belonging to the order are almost invariably included in the collections brought back by members of Arctic expeditions. Others are met with in the most isolated localities; thus the Rev. A. E. Eaton discovered on the desolate shores of Kerguelen’s Island apterous and semi-apterous Diptera (Tipulidae and Ephydridae) of a degraded type adapted to the climatic peculiarities of the locality. Many bird parasites belonging to the Hippoboscidae have naturally been carried about the world by their hosts, while other species, such as the house-fly, blow-fly and drone-fly, have in like manner been disseminated by human agency. Most families and a large proportion of genera are represented throughout the world, but in some cases (e.g. Glossina—see Tsetse-Fly) the distribution of a genus is limited to a continent. As a rule the general facies as well as dimensions are remarkably uniform throughout a family, so that tropical species often differ little in appearance from those inhabiting temperate regions. Many instances of exaggerated and apparently unnatural structure nevertheless occur, as in the case of the genera Pangonia, Nemestrina, Achias, Diopsis and the family Celyphidae, and, as might be expected, it is chiefly in tropical species that these peculiarities are found. To a geographical distribution of the widest extent, Diptera add a range of habits of the most diversified nature; they are both animal and vegetable feeders, an enormous number of species acting, especially in the larval state, as scavengers in consuming putrescent or decomposing matter of both kinds. The phytophagous species are attached to various parts of plants, dead or alive; and the carnivorous in like manner feed on dead or living flesh, or its products, many larvae being parasitic on living animals of various classes (in Australia the larva of a species of Muscidae is even a parasite of frogs), especially the caterpillars of Lepidoptera, which are destroyed in great numbers by Tachininae. The recent discovery of a bloodsucking maggot, which is found in native huts throughout the greater part of tropical and subtropical Africa, and attacks the inmates when asleep, is of great interest.
It may confidently be asserted that, of insects which directly or indirectly affect the welfare of man, Diptera form the vast majority, and it is a moot point whether the good effected by many species in the rapid clearing away of animal and vegetable impurities, and in keeping other insect enemies in check, counterbalances the evil and annoyance wrought by a large section of the Order. The part played by certain blood-sucking Diptera in the dissemination of disease is now well known (see Mosquito and Tsetse-Fly), and under the term myiasis medical literature includes a lengthy recital of instances of the presence of Dipterous larvae in various parts of the living human body, and the injuries caused thereby. That Diptera of the type of the common house-fly are often in large measure responsible for the spread of such diseases as cholera and enteric fever is undeniable, and as regards blood-sucking forms, in addition to those to which reference has already been made, it is sufficient to mention the vast army of pests constituted by the midges, sand-flies, horse-flies, &c., from the attacks of which domestic animals suffer equally with man, in addition to being frequently infested with the larvae of the bot and warble flies (Gastrophilus, Oestrus and Hypoderma). Lastly, as regards the phytophagous forms, there can be no doubt that the destruction of grass-lands by “leather-jackets” (the larvae of crane-flies, or daddy-long-legs,—Tipula oleracea and T. paludosa), of divers fruits by Ceratitis capitata and species of Dacus, and of wheat and other crops by the Hessian-fly (Mayetiola destructor) and species of Oscinis, Chlorops, &c., is of very serious consequence.
With many writers it is customary to treat the fleas as a sub-order of Diptera, under the title Aphaniptera or Siphonaptera. Since, however, although undoubtedly allied to the Diptera, they must have diverged from the ancestral stem at an early period, before the existing forms of Diptera became so extremely specialized, it seems better to regard the fleas as constituting an independent order (see Flea). (E. E. A.)
DIPTERAL (Gr. for “double-winged”), the architectural term applied to those temples which have a double range of columns in the peristyle, as in the temple of Diana at Ephesus.
DIPTYCH (Gr. δίπτυχος, two-folding), (1) A tablet made with a hinge to open and shut, used in the Roman empire for letters (especially love-letters), and official tokens of the commencement of a consul’s, praetor’s or aedile’s term of office. The latter variety of diptych was inscribed with the magistrate’s name and bore his portrait, and was issued to his friends and the public generally. They were made of boxwood or maple. More costly examples were in cedar, ivory (q.v.), silver or sometimes gold. They were often sent as New Year gifts.
(2) In the primitive church when the worshippers brought their own offerings of bread and wine, from which were taken the Communion elements, the names of the contributors were recorded on diptychs and read aloud. To these names were early added those of deceased members of the community whom it was desired to commemorate. This custom rapidly developed into a kind of commemoration of saints and benefactors, living and dead; especially, in each church, were the names of those who had been its bishops recorded. The custom was maintained until the lists became so long that it was impossible to read them through, and the observance in this form had to be abandoned. The insertion of a name on the diptych, thereby securing the prayers of the church, was a privilege from which a person could be excluded on account of suspicion of heresy or by the intrigues of enemies. His name could, if written, be expunged under similar circumstances. The names thus written were read from
