circumstances. From peculiarities in the examples of Acridium peregrinum taken in England in 1869, it has been asserted that they must have come direct by sea from the west coast of Africa; and what is probably the same species has been seen in the Atlantic at least 1200 m. from land, in swarms completely covering the ship; thus, in certain cases flight must be sustained for several days and nights together. The height at which swarms fly, when their horizontal course is not liable to be altered by mountains, has been very variously estimated at from 40 to 200 ft., or even in a particular case to 500 ft. The extent of swarms and the number of individuals in a swarm cannot be accurately ascertained. They come sometimes in such numbers as to completely obscure the sun, when the noise made by the rustling of the wings is deafening. Nevertheless some idea on this point may be formed from the ascertained fact that in Cyprus in 1881, at the close of the season, 1,600,000,000 egg-cases, each containing a considerable number of eggs, had been destroyed; the estimated weight exceeding 1300 tons. Yet two years later, it is believed that not fewer than 5,076,000,000 egg-cases were again deposited in the island.
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Fig. 1.—Pachytylus migratorius. This and the other figures are all natural size. |
In Europe the best known and ordinarily most destructive species is Pachytylus cinerascens, and it is to it that most of the numerous records of devastations in Europe mainly refer, but it is probably not less destructive in many parts of Africa and Asia. That the arid steppes of central Asia are the home of this insect appears probable; still much on this point is enveloped in uncertainty. In any case the area of permanent distribution is enormous, and that of occasional distribution is still greater. The former area extends from the parallel of 40° N. in Portugal, rising to 48° in France and Switzerland, and passing into Russia at 55°, thence continuing across the middle of Siberia, north of China to Japan; thence south to the Fiji Islands, to New Zealand and North Australia; thence again to Mauritius and over all Africa to Madeira. The southern distribution is uncertain and obscure. Taking exceptional distribution, it is well known that it occasionally appears in the British Isles, and has in them apparently been noticed as far north as Edinburgh; so also does it occasionally appear in Scandinavia, and it has probably been seen up to 63° N. in Finland. Looking at this vast area, it is easy to conceive that an element of uncertainty must always exist with regard to the exact determination of the species, and in Europe especially is this the case, because there exists a distinct species, known as P. migratorius, the migratory area of which appears to be confined to Turkestan and eastern Europe.
P. cinerascens is certainly the most common of the “locusts” occasionally found in the British Isles, and E. de Selys-Longchamps is of opinion that it breeds regularly in Belgium, whereas the true P. migratorius is only accidental in that country.
Fig. 2.—Acridium peregrinum.
A South African species allied to the preceding and provisionally identified as Pachytylus salcicollis is noteworthy from the manifestation of the migratory instinct in immature wingless individuals. The families of young, after destroying the vegetation of a district, unite in a vast army and move away in search of fresh pastures, devastating the country as they go and proceeding of necessity on foot, hence they are known to the Dutch as “voetgangers.” Travelling northwards towards the centre of the continent, the home of their parents before migration, they are diverted from their course by no obstacles. Upon reaching a river or stream they search the bank for a likely spot to cross, then fearlessly cast themselves upon the water where they form floating islands of insects, most of which usually succeed in gaining the opposite bank, though many perish in the attempt.
Acridium peregrinum (fig. 2) can scarcely be considered even an accidental visitor to Europe; yet it has been seen in the south of Spain, and in many examples spread over a large part of England in the year 1869. It is a larger insect than P. migratorius. There is every reason to believe that it is the most destructive locust throughout Africa and in India and other parts of tropical Asia, and its ravages are as great as those of P. migratorius. Presumably it is the species occasionally noticed in a vast swarm in the Atlantic, very far from land, and presumably also it occurs in the West Indies and some parts of Central America. In the Argentine Republic a (possibly) distinct species (A. paranense) is the migratory locust.
Caloptenus italicus (fig. 3) is a smaller insect, with a less extended area of migration; the destruction occasioned in the districts to which it is limited is often scarce less than that of its more terrible allies. It is essentially a species of the Mediterranean district, and especially of the European side of that sea, yet it is also found in North Africa, and appears to extend far into southern Russia.
Caloptenus spretus (fig. 4) is the “Rocky Mountain locust” or “hateful grasshopper” of the North American continent. Though a comparatively small insect, not so large as some of the grasshoppers of English fields, its destructiveness has procured for it great notoriety. By early travellers and settlers the species was not recognized as distinct from some of its non-migratory congeners. But in 1877, Congress appointed a United States Entomological Commission to investigate the subject. The report of the commissioners (C. V. Riley, A. S. Packard and C. Thomas) deals with the whole subject of locusts both in America and the Old world. C. spretus has its home or permanent area in the arid plains of the central region east of the Rocky Mountains, extending slightly into the southern portion of Canada; outside this is a wide fringe to which the term sub-permanent is applied, and this is again bounded by the limits of only occasional distribution, the whole occupying a large portion of the North American continent; but it is not known to have crossed the
