This series of feet represents the evolutionary succession
from the Eocene Hypohippus (1) to the modern Equus (6)
seen in front and in side view. The top bone is the os calcis,
or hock bone, to which the tendon Achilles is attached. The
bottom bone is the terminal phalanx which is inserted in the
heart of the hoof.
Equus caballus.
Modern horse.
Merychippus sp.
Miocene.
Merychippus insignis (milk molar).
Parahippus pawniensis.
Mesohippus intermedius.
Upper Oligocene (White river formation).
Mesohippus bairdi?
Oligocene (White river formation).
Mesohippus bairdi.
Orohippus sp.
Middle Eocene (Bridger formation).
Eohippus sp.
Lower Eocene (Wind river formation). (Wasatch formation).
Eohippus sp.
1
2
3
4
5
6
The stages are as follows:
1. Hypohippus, Lower Eocene.
4. Protohippus, Upper Miocene.
2. Mesohippus, Lower Oligocene.
5. Neohipparion, Upper Miocene.
3. Parahippus, Lower Miocene.
6. Equus, Pleistocene and recent.
The evolution consists first in progressive increase
in size; second, in the acceleration of the
median digit and retardation of the lateral digits,
the latter becoming more and more elevated from
the ground until finally in Equus (6) they are the
lateral splints, which in the embryonic condition
have vestigial cartilages attached
representing the last traces of the
lateral phalanges.
1
2
3
4
5
6
Fig. 8.—TEN STAGES IN THE EVOLUTION
OF THE SECOND UPPER
MOLAR TOOTH OF THE RIGHT
SIDE, ARRANGED ACCORDING TO
GEOLOGICAL LEVEL.
(Nos. 1-9 from “American Equidae.”)
Fig. 7.—LAW OF ACCELERATION AND RETARDATION ILLUSTRATED IN
THE EVOLUTION OF THE HIND FEET OF THE HORSE.
(From photos lent by the American Museum of Natural History.)