The great majority of botanists in the second half of the
18th century had no longer any doubt that the stamens were
organs of reproduction, and they were anxious to prove the
existence of similar organs in the Cryptogams; they rested in
this matter on external resemblances and analogies, which they
interpreted in a more or less arbitrary manner. The obvious
external resemblance between the antheridia and archegonia in
Mosses and the sexual organs in the Phanerogams led Schmidel
and Hedwig to consider them to be stamens and ovaries, and
the conjecture was correct, though the true nature of the moss-fruit had to be learnt in another way. Micheli, Linnaeus
and Dillen, trusting still more to external appearance and with
slight knowledge of these plants, had before this taken the
fruit for a male flower, and in the case of the rest of the
Cryptogams the best botanists were only feeling their way in
the dark with no certain experience to guide them. It is not
necessary to give a particular account of the views which
originated in this way; one or two may be mentioned by way
of example. Koelreuter regarded the volva of Mushrooms,
Gleditsch and Hedwig certain tube-like cells in their lamellae,
as the male organs of fertilisation. Gleichen took the stomata,
Koelreuter the indtisium, Hedwig even the glandular hairs of
Ferns for anthers. It was not yet suspected that the course of
development and the whole morphology of the Cryptogams
could not be so compared with that of the Phanerogams;
correct and incorrect assumptions with regard to the sexual
organs of the Cryptogams were alike devoid of scientific value,
being mere guesses and vague conjectures. Nor was the state
of things much better even in the first years of the 19th century; and if by that time a number of occasional observations had been made which could afterwards be turned to
scientific account, these were as yet only isolated facts without
scientific connection, and every one was at liberty to concede
or to refuse sexual organs to the Cryptogams generally at his own discretion. Meanwhile observations gradually accumu-
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Appearance
Chap. I.]
Sexuality in Cryptogams.
437