Page:Makers of British botany.djvu/337

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LOGARITHMIC CURVE OF GROWTH
273

Comparatively little was known of the life history of any Schizomycete. Ward therefore made a detailed and exhaustive study of that of Bacillus ramosus, the Wurzel bacillus of German authors, which is common in Thames water, and bears a superficial resemblance to the anthrax bacillus, but is innocuous. It proved convenient for study, as it ran through its entire life history in from thirty to sixty hours at ordinary temperature. It forms long filaments, the growth of which Ward was able to measure under the microscope with great precision. On plotting out his measurements he obtained a regular curve, from which he found that, under constant conditions, the filament doubled itself in equal times. This he called "the law of doubling." It is the same as the so-called "law of compound interest," and leads to the expression of the growing quantity as an exponential function of the time, so that the time is proportional to the logarithm of that quantity. This relation has, of course, long been familiar in chemical reactions, but, as far as I know, Ward was the first to detect it in any vital process in a plant. This, which was in 1895, has, I think, been overlooked. Stefanowska has since, in 1904, obtained a logarithmic curve for the early period of the growth of maize, which doubles its weight every ten days, and the subject has since been pursued by Chodat and others.

In speculating on the cause of the destructive action of light on bacteria, Ward adopted the view of his friend Elfving, that it inhibited metabolic processes necessary to nutrition. He suggests that the "constructed metabolites" at the moment of assimilation are in a highly unstable condition, and liable to destruction by oxidation promoted by light. He points to the fact that plant structures are frequently provided with colour screens, which would cut off the blue-violet rays and check their action in promoting the rapid oxidation of reserve materials, and he quotes the suggestion of Elfving that chlorophyll itself may serve as such a screen against "destructive metabolic action in synthesis." Ward seems to have attributed little importance to the fact that substantially the same view had long before been put forward by Pringsheim, though received with little favour. His own view that when red and orange predominate in the