collection has yielded evidence of the most striking character in regard to this question of the. pedigree of the horse. Indeed, the evidence which Prof. Marsh has collected tends to show that you have in America the true original seat of the equine type—the country in which the evidence of the primitive form and successive modifications of the horse series is far better preserved than in Europe; and Prof. Marsh's kindness has enabled me to put before you the following diagram, every figure in which is an actual representation of the specimen which is preserved in New Haven at this present time. The succession of forms which he has brought together shows, in the first place, the great care and patience to which I have referred. Secondly, there is this Pliocene form of the horse (Pliohippus); the conformation of its limbs presents some very slight deviations from the ordinary horse, and with shorter crown of the grinding teeth. Then comes the form which represents the European Hipparion, which is the Protohippus, having three toes and the forearm and leg and teeth to which I have referred, and which is more valuable than the European Hipparion for this reason: it is devoid of some of the peculiarities of that form—peculiarities which tend to show that the European Hipparion is rather a side branch than one in the direct line of succession. Next comes the Miohippus, which corresponds pretty nearly with what I spoke of as the Anchitherium of Europe, but which has some interesting peculiarities. It presents three toes—one large median and two lateral ones; of the toe which answers to the little finger of the human hand, there is only a rudiment. This is, however, as far as European deposits have been enabled to carry us with any degree of certainty in the history of the horse. In the American tertiaries, on the contrary, the series of equine forms is continued down to the bottom of the Eocene. The older Miocene form, termed Mesohippus, has three toes in front and a large splint-like rudiment representing the little finger, and three toes behind. The radius and ulna are entire, and the tibia and fibula distinct, and the teeth are anchitheroid with short crowns.
But the most important discovery of all is the Orohippus—which comes from the lower part of the Eocene formation, and is the oldest member of the equine series known. Here we have four complete toes on the front-limb, three toes on the hind-limb, a well-developed ulna, a well-developed fibula, and the teeth of simple pattern. So you are able, thanks to these great researches, to show that, so far as present knowledge extends, the history of the horse-type is exactly and precisely that which could have been predicted from a knowledge of the principles of evolution. And the knowledge we now possess justifies us completely in the anticipation that when the still lower Eocene' deposits and those which belong to the Cretaceous epoch have yielded up their remains of equine animals, we shall find first an equine creature with four complete toes and a rudiment of the innermost toe in front, and probably a rudiment of the fifth toe in the
