that the latter is prepared, even before individual experience, to perform adjustive actions mechanically which, in previous generations, were performed intelligently. This mode of origin of instincts has been called by Mr. Lewes the "lapsing of intelligence," and it was fully recognized by Mr. Darwin as a factor in the formation of instinct.
The Darwinian theory of instinct, then, attributes the evolution of instincts to these two causes acting either singly or in combination—natural selection and lapsing intelligence. I shall now proceed to adduce some of the more important facts and considerations which, to the best of my judgment, support this theory, and show it to be by far the most comprehensive and satisfactory explanation of the phenomena which has hitherto been propounded.
That many instincts must have owed their origin and development to natural selection exclusively is, I think, rendered evident by the following general considerations:
1. Considering the great importance of instincts to species, we are prepared to expect that they must be in large part subject to the influence of natural selection. 2. Many instinctive actions are performed by animals too low in the scale to admit of our supposing that the adjustments which are now instinctive can ever have been intelligent. 3. Among the higher animals instinctive actions are performed at an age before intelligence, or the power of learning by individual experience, has begun to assert itself 4. Many instincts, as we now find them, are of a kind which, although performed by intelligent animals at a matured age, yet can obviously never have been originated by intelligent observation. Take, for instance, the instinct of incubation. It is quite impossible that any animal, prior to individual or ancestral experience, can have kept its eggs warm with the intelligent purpose of developing their contents; so we can only suppose that the incubating instinct began in some such form as we now see it in the spider, where the object of the process is protection, as distinguished from the imparting of heat. But incidental to such protection in the case of a warm-blooded animal is the imparting of heat, and, as animals gradually became warm-blooded, no doubt this latter function became of more and more importance to incubation. Consequently, those individuals which most constantly cuddled their eggs would develop most progeny, and so the incubating instinct would be developed by natural selection without there ever having been any intelligence in the matter.
From those four general considerations, therefore, we may conclude (without waiting to give special illustrations of each) that one mode of origin of instincts consists in natural selection, or survival of the fittest, continuously preserving actions which, although never intelligent, yet happen to have been of benefit to the animals which first chanced to perform them. Among animals, both in a state of nature and domes-
