separates the digestive cavity from the outer sea-water, and that is drawn out in processes to form the hollow tentacles. In no part of its structure is the sea-anemone massive, as is the case in most higher forms, where the muscles, skeleton and so forth usually give rise to a considerable thickness of tissue; in fact, the animal exhibits no welldefined organs except the digestive organs, and may be described as a membranous digestive sac.
Although the body of the sea-anemone is really nothing more than membranous walls, these walls have long been known to contain both nerve and muscle. These two tissues occur over almost the whole animal. According to the Hertwigs, the nervous tissue is more abundant in the neighborhood of the mouth than elsewhere, and this region has been regarded by some investigators as a central nervous organ. But the studies of Jordan and others have shown conclusively that this opinion is not correct, and that the removal of this region interferes in no serious way with the reactions of the animal. Apparently each part of the sea-anemone carries with it its own neuromuscular mechanism, a condition well illustrated by the tentacles. These organs are chiefly concerned with appropriating the food and are stimulated by the dissolved materials in the food. A tentacle when cut off from a sea-anemone and held in sea-water can still be stimulated by food and will exhibit almost exactly the same kind of movements when thus isolated that it did when a part of the whole animal, thus demonstrating the completeness and independence of its own neuromuscular mechanism. Nervous transmission can be accomplished from almost any part of the sea-anemone to almost any other part, but as such experiments as those with the tentacles indicate, no one part of the animal's nervous organization seems to be more important than any other part. In other words, the nervous system in the sea-anemone is diffuse rather than centralized.
When the minute organization of the nervous system of these animals is studied, it is found to consist of a vast number of sensory neurones which connect the surface of the animal with the underlying muscles and which form there what appears to be an intricate nervous network. This nervous mechanism is concerned primarily with the reception of stimuli and the immediate excitation of the muscles. The nervous mechanism is a receptor mechanism that acts as a trigger for setting off the muscle. The whole neuromuscular apparatus seems to be made up of those two elements which in the higher animals were designated receptors and effectors and without the intervention of an adjustor or central nervous organ. Viewed from the standpoint of development, this condition points indubitably to the conclusion that the central nervous organs were evolved only after the appearance of sense organs and muscle, and that such animals as the sea-anemone may well be taken
