I conceive that distinct provinces of the distribution of terrestrial life have existed since the earliest period at which that life is recorded, and possibly much earlier; and I suppose, with Mr. Darwin, that the progress of modification of terrestrial forms is more rapid in areas of elevation than in areas of depression. I take it to be certain that Labyrinthodont Amphibia existed in the distributional province which included the dry land depressed during the Carboniferous epoch; and I conceive that, in some other distributional provinces of that day, which remained in the condition of stationary or of increasing dry land, the various types of the terrestrial Sauropsida and of the Mammalia were gradually developing.
The Permian epoch marks the commencement of a new movement of upheaval in our area, which attained its maximum in the Triassic epoch, when dry land existed in North America, Europe, Asia, and Africa, as it does now. Into this great new continental area the Mammals, Birds, and Reptiles developed during the Palæozoic epoch spread, and formed the great Triassic Arctogæal province. But, at the end of the Triassic period, the movement of depression recommenced in our area, though it was doubtless balanced by elevation elsewhere; modification and development, checked in the one province, went on in that "elsewhere"; and the chief forms of Mammals, Birds, and Reptiles, as we know them, were evolved and peopled the Mesozoic continent. I conceive Australia to have become separated from the continent as early as the end of the Triassic epoch, or not much later. The Mesozoic continent must, I conceive, have lain to the east, about the shores of the North Pacific and Indian Oceans; and I am inclined to believe that it continued along the eastern side of the Pacific area to what is now the province of Austro-Columbia, the characteristic Fauna of which is probably a remnant of the population of the latter part of this period.
Towards the latter part of the Mesozoic period the movement of upheaval around the shores of the Atlantic once more recommenced, and was very probably accompanied by a depression around those of the Pacific. The Vertebrate fauna elaborated in the Mesozoic continent moved westward and took possession of the new lands, which gradually increased in extent up to, and in some directions after, the Miocene epoch.
It is in favour of this hypothesis, I think, that it is consistent with the persistence of a general uniformity in the positions of the great masses of land and water. From the Devonian period, or earlier, to the present day, the four great oceans, Atlantic, Pacific, Arctic, and Antarctic, may have occupied their present positions, and only their coasts and channels of communication have undergone an incessant alteration. And, finally, the hypothesis I have put before you requires no supposition that the rate of change in organic life has been either greater or less in ancient times than it is now; nor any assumption, either physical or biological, which has not its justification in analogous phenomena of existing nature.
I have now only to discharge the last duty of my office, which is
