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ARDEA MEGACEPHALAMILNE-EDWARDS.
- Butors Leguat, Relation du Voyage (1708).
- Ardea megacephala Milne-Edwards, Ann. Sci. Nat. (5) XIX, 1874, p. 10.
Leguat's description, here translated, is as follows:—"We had Bitterns as big and as fat as capons. They are tamer and more easily caught than the 'gelinotes.'" He also says, "The lizards often serve as prey for the birds, especially for the Bitterns. When we shook them down from the branches with a pole, these birds ran up and gobbled them down in front of us, in spite of all we could do to prevent them; and even if we only pretended to do so they came in the same manner and always followed us about."
Milne-Edwards remarks, among other notes, that "This bird is not a true Bittern, but its head is so large and its feet so short that it is easy to understand that Leguat should have called it so."
The bony structure of the head is remarkable on account of its massive and thick proportions; the skull itself is strongly enlarged posteriorly, and the temporal fossae are bordered by very pronounced ridges, especially those on the occipital region. The upper side of the skull is hardly convex, and the interorbital region is large, but only slightly depressed along its middle line. The bill is stout, almost straight, a good deal enlarged at its base and rounded beneath. The nostrils are large and preceded by a large groove, which extends very far towards the tip.
It is impossible to confound this skull with that of any Bittern, the latter having the beak relatively slender and only barely exceeding the skull in length. These also have the skull much constricted at the temporal region. The fossil skull from Rodriguez therefore presents characters essentially those of a Heron, but differs from all known species in its massive appearance. In the Grey, Purple and Goliath Herons, as well as in the Egrettes, the head is narrower, more elongated, the bill less conical and less strong. In Ardea atricollis, now inhabiting Madagascar, the beak much resembles that of our extinct species, but it is longer and less enlarged at the base. The interorbital area is much wider, while on the other hand the hinder portion of the skull is narrower and more elongated, which gives to the skull a totally different aspect.
The feet relatively to the head are extremely short, and from this I conclude that we know no species of Heron which can be compared to that of Rodriguez. Nevertheless, the tarso-metatarsus presents all the characters
