(b) Movements and Temperature of Lake-Waters.—(1) In addition to the rise and fall of the surface-level of lakes due to rainfall and evaporation, there is a transference of water due to the action of wind which results in raising the level at the end to which the wind is blowing. In addition to the well-known progressive waves there are also stationary waves or “seiches” which are less apparent. A seiche is a standing oscillation of a lake, usually in the direction of the longest diameter, but occasionally transverse. In a motion of this kind every particle of the water of the lake oscillates synchronously with every other, the periods and phases being the same for all, and the orbits similar but of different dimensions and not similarly situated. Seiches were first discovered in 1730 by Fatio de Duillier, a well-known Swiss engineer, and were first systematically studied by Professor Forel in the Lake of Geneva. Large numbers of observations have been made by various observers in lakes in many parts of the world. Henry observed a fifteen-hour seiche in Lake Erie, which is 396 kilometres in length, and Endros recorded a seiche of fourteen seconds in a small pond only 111 metres in length. Although these waves cause periodical rising and falling of the water-level, they are generally inconspicuous, and can only be recorded by a registering apparatus, a limnograph. Standard work has been done in the study of seiches by the Lake Survey of Scotland under the immediate direction of Professor Chrystal, who has given much attention to the hydrodynamical theories of the phenomenon. Seiches are probably due to several factors acting together or separately, such as sudden variations of atmospheric pressure, changes in the strength or direction of the wind. Explanations such as lunar attraction and earthquakes have been shown to be untenable as a general cause of seiches.
2. The water temperature of lakes may change with the season from place to place and from layer to layer; these changes are brought about by insolation, by terrestrial radiation, by contract with the atmosphere, by rain, by the inflow of rivers and other factors, but the most important of all these are insolation and terrestrial radiation. Fresh water has its greatest density at a temperature of 39.2° F., so that water both above and below this temperature floats to the surface, and this physical fact largely determines the water stratification in a lake. In salt lakes the maximum density point is much lower, and does not come into play. In the tropical type of fresh-water lake the temperature is always higher than 39° F., and the temperature decreases as the depth increases. In the polar type the temperature is always lower than 39° F., and the temperature increases from the surface downwards. In the temperate type the distribution of temperature in winter resembles the polar type, and in summer the tropical type. In Loch Ness and other deep Scottish lochs the temperature in March and April is 41° to 42° F., and is then nearly uniform from top to bottom. As the sun comes north, and the mean air temperature begins to be higher than the surface temperature, the surface waters gain heat, and this heating goes on till the month of August. About this time the mean air temperature falls below the surface temperature, and the loch begins to part with its heat by radiation and conduction. The temperature of the deeper layers beyond 300 ft. is only slightly affected throughout the whole year. In the autumn the waters of the loch are divided into two compartments, the upper having a temperature from 49° to 55° F., the deeper a temperature from 41° to 45°. Between these lies the discontinuity-layer (Sprungschicht of the Germans), where there is a rapid fall of temperature within a very short distance. In August this discontinuity-layer is well marked, and lies at a depth of about 150 ft.; as the season advances this layer gradually sinks deeper, and the layer of uniform temperature above it increases in depth, and slowly loses heat, until finally the whole loch assumes a nearly uniform temperature. Many years ago Sir John Murray showed by means of temperature observations the manner in which large bodies of water were transferred from the windward to the leeward end of a loch, and subsequent observations seem to show that, before the discontinuity-layer makes its appearance, the currents produced by winds are distributed through the whole mass of the loch. When, however, this layer appears, the loch is divided into two current-systems, as shown in the following diagram:—
 | |
| Current systems in a loch induced by wind at the surface. (After Wedderburn.) | |
AB, Discontinuity layer. |
E, Secondary surface current. |
Another effect of the separation of the loch into two compartments by the surface of discontinuity is to render possible the temperature-seiche. The surface-current produced by the wind transfers a large quantity of warm water to the lee end of the loch, with the result that the surface of discontinuity is deeper at the lee than at the windward end. When the wind ceases, a temperature-seiche is started, just as an ordinary seiche is started in a basin of water which has been tilted. This temperature-seiche has been studied experimentally and rendered visible by superimposing a layer of paraffin on a layer of water.
Wedderburn estimates the quantity of heat that enters Loch Ness and is given out again during the year to be approximately sufficient to raise about 30,000 million gallons of water from freezing-point to boiling-point. Lakes thus modify the climate of the region in which they occur, both by increasing its humidity and by decreasing its range of temperature. They cool and moisten the atmosphere by evaporation during summer, and when they freeze in winter a vast amount of latent heat is liberated, and moderates the fall of temperature.
Lakes act as reservoirs for water, and so tend to restrain floods, and to promote regularity of flow. They become sources of mechanical power, and as their waters are purified by allowing the sediment which enters them to settle, they become valuable sources of water-supply for towns and cities. In temperate regions small and shallow lakes are likely to freeze all over in winter, but deep lakes in similar regions do not generally freeze, owing to the fact that the low temperature of the air does not continue long enough to cool down the entire body of water to the maximum density point. Deep lakes are thus the best sources of water-supply for cities, for in summer they supply relatively cool water and in winter relatively warm water. Besides, the number of organisms in deep lakes is less than in small shallow lakes, in which there is a much higher temperature in summer, and consequently much greater organic growth. The deposits, which are formed along the shores and on the floors of lakes, depend on the geological structure and nature of the adjacent shores.
Biology.—Compared with the waters of the ocean those of lakes may safely be said to contain relatively few animals and plants. Whole groups of organisms—the Echinoderms, for instance—are unrepresented. In the oceans there is a much greater uniformity in the physical and chemical conditions than obtains in lakes. In lakes the temperature varies widely. To underground lakes light does not penetrate, and in these some of the organisms may be blind, for example, the blind crayfish (Cambarus pellucidus) and the blind fish (Amblyopsis spelaeus) of the Kentucky caves. The majority of lakes are fresh, while some are so salt that no organisms have been found in them. The peaty matter in other lakes is so abundant that light does not penetrate to any great depth, and the humic acids in solution prevent the development of some species. Indeed, every lake has an individuality of its own, depending upon climate, size, nature of the bottom, chemical composition and connexion with other lakes. While the ocean contains many families and genera not represented in lakes, almost every genus in lakes is represented in the ocean.
The vertebrates, insects and flowering plants inhabiting lakes vary much according to latitude, and are comparatively well known to zoologists and botanists. The micro-fauna and flora have only recently been studied in detail, and we cannot yet be said to know much about tropical lakes in this respect. Mr James Murray, who has studied the Scottish lakes, records in over 400 Scottish lochs 724 species (the fauna including 447 species, all invertebrates, and the flora comprising 277 species) belonging to the following groups; the list must not be regarded as in any way complete:—
| Fauna. | Flora. | ||||
| Mollusca | 7 | species | Phanerogamia | 65 | species |
| Hydrachnida | 17 | ” | Equisetaceae | 1 | ” |
| Tardigrada | 30 | ” | Selaginellaceae | 1 | ” |
| Insecta | 7 | ” | Characeae | 6 | ” |
| Crustacea | 78 | ” | Musci | 18 | ” |
| Bryozoa | 7 | ” | Hepaticae | 2 | ” |
| Worms | 25 | ” | Florideae | 2 | ” |
| Rotifera | 181 | ” | Chlorophyceae | 142 | ” |
| Gastrotricha | 2 | ” | Bacillariaceae | 26 | ” |
| Coelenterata | 1 | ” | Myxophyceae | 10 | ” |
| Porifera | 1 | ” | Peridiniaceae | 4 | ” |
| Protozoa | 91 | ” | |||
| —————— | —————— | ||||
| 447 | ” | 277 | ” | ||
These organisms are found along the shores, in the deep waters, and in the surface waters of the lakes.
The littoral region is the most populous part of lakes; the existence of a rooted vegetation is only possible there, and this in turn supports a rich littoral fauna. The greater heat of the water along the margins also favours growth. The great majority of the species in Scottish lochs are met with in this region. Insect larvae of many kinds are found under stones or among weeds. Most of the Cladocera, and the
