Page:Plant succession; an analysis of the development of vegetation.pdf/10

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CONTENTS.

VI

III. Initial Causes - Continued


Climatic Causes

Róle 55
Bare areas due to climatic factors directly 55
Bare areas due to drouth 56
Bare areas due to wind 56
Bare areas due to snow, hail, and frost 56
Bare areas due to lightning 57
Bare areas due indirectly to climatic factors 57
Sudden changes of climate 57

Biotic Causes

General relations 58
Action and effect 58
Bare areas due to destruction of vegetation alone 59
Bare areas with dry or drier soils 59
Bare areas with wet soils or water 60

Primary and Secondary Areas

Distinction 60
Sterility of primary and secondary areas 60
Denudation 61
Methods of denudation 61
Depth of removal or deposit 61
Rate and extent of removal 62

Ecesic Causes

Nature 63

Aggregation

Concept and róle 63
Effects of simple aggregation 63
Relation to denuded areas 64
Interaction of aggregation and migration 64

Migration

Concept 64
Mobility 64
Seed-production 65
Influence of the organ used 65
Influence of the migration contrivance 66
Role of migration agents 67
Destructive action of agents 67
Direction of migration 67

Ecesis

Nature and róle 68
Germination 69
Fate of seedling 70
Growth 71
Reproduction 71
Ecesis in bare areas 71

Competition

Nature 72
Competition and dominance 72
Competition in air and in soil 73
Róle of competition in succession 73

Invasion

Nature and róle 75
Kinds of invasion 75
Manner of invasion 76
Barriers 77
Biological barriers 77
Changes in barriers 78

Reactions

Concept and nature 79
Róle in succession 80
Previous analyses of reaction 80
Kinds of reactions 81

Soil Formation

Manner 81
Reaction by accumulating plant bodies or parts 81
Reaction by accumulating plant concretions 83
Reaction by producing weathering 83
Reaction upon wind-borne material 84
Reaction upon water-borne detritus 85
Reaction upon slipping sand and gravel 86

Soil Structure

Reaction by adding humus 86
Reaction by compacting the soil 87
Reaction by preventing weathering or erosion 88

Water-Content

Reaction by increasing water-content 88
Reaction by decreasing water-content 89

Nutrients and Solutes

Reaction by adding nutrients or foodstuffs 89
Reaction by decreasing nutrients 89
Reaction by producing acids 90
Reaction by producing toxins 90

Soil Organisms

Reaction by means of parasites 91
Reaction by means of saprophytes 92

Air Reactions

Reaction upon light 92
Reaction upon humidity, temperature, and wind 94
Reaction upon local climate 94
Reaction upon aërial organisms 95
Correlation of reactions 96
Quantitative study of reactions 96

Stabilization and Climax

Stabilization 98
Causes of stabilization 98
Relation to the climax 98
Degree of stabilization 99

Life-History Stages

Nature
Kinds of stages 100
Role of life forms 100
Reasons why plants disappear 100
Reasons why plants appear at certain stages 100
Reasons why plants appear before their proper time 103
Initial stages 103
Medial stages 105